The British Rail Class 421 (4CIG) electric multiple units were built at BR's Holgate Road carriage works between 1964 and 1972. Units were built in two batches, and were initially introduced on services on the Brighton Main Line. Later units were introduced on services to Portsmouth. These units replaced older Southern Railway-designed units, such as the 5Bel "Brighton Belle" units, and 4Cor units. The fleet's lifespan was 46 years.