In biology, paternal care is parental investment provided by a male to his own offspring. It is a complex social behaviour in vertebrates associated with animal mating systems, life history traits, and ecology. Paternal care may be provided in concert with the mother (biparental care) or, more rarely, by the male alone (so called exclusive paternal care). Exclusive paternal care has evolved multiple times in a variety of organisms, including invertebrates, fishes, and amphibians.