The evolution of fungi has been going on since fungi diverged from other life around 1.5 billion years ago, with the glomaleans branching from the "higher fungi" (dikaryans) at ~570 million years ago, according to DNA analysis. (Schüssler et al., 2001; Tehler et al., 2000) Fungi probably colonized the land during the Cambrian, over 500 million years ago, (Taylor & Osborn, 1996), and possibly 635 million years ago during the Ediacaran, but terrestrial fossils only become uncontroversial and common during the Devonian, 400 million years ago.